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pDH51 can be used to express gene sequences in plant cells; contains the promoter of the cauliflower mosaic virus (CaMV) 35S RNA gene and its transcription terminator, separated by a polylinker sequence with several unique restriction sites; for direct plant protoplast transformation or can be adapted as part of the Ti transformation system; the presence of the lac promoter allows expression in Escherichia coli of genes inserted into the polylinker sequence.